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Long J, Liu XK, Kang ZP, Wang MX, Zhao HM, Huang JQ, et al

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It undergoes hydrolysis by intracellular esterase, releasing Fluo-4, which binds to calcium with a K d value of 345 nM and emits green fluorescence [45]

So instead of one obvious interaction, you get a bunch of subtle ones layered together

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